I'd say the fact that they have the black around their Sharingan eyes, cracked skin, and we SAW Kabuto summon them from coffins makes it a pretty safe bet that they're Edo Tensei reanimations.
Plus, we've seen the Two Tails chick regenerate already. Nagato's Pains would only regenerate if he threw them inside the demon head.
I think they're obviously reanimated by Kabuto, but Tobi is using the Chakra receivers to control them, share his eyesight with them, and infuse them with Bijuu chakra.
They did, and Kakashi thinks these should have those as well, but don't because controlling the Bijuu is taking up too much of his concentration(which, frankly, just feel like a lazy writing excuse because Kishi has given too many powers to this character)